Drej Queen Susquehana

Drej Queen Susquehana is the main antagonist of Titan A.E.. She is the ruler of an energy-based alien species called the Drej. She is the only female Drej and the only member of the species to have eyes while her followers are eyeless drones. She intends to get rid of all humanity due to an experiment called the Titan Project. In the film she speaks an alien language with English subtitles. Although she is a female, she was voiced by Christopher Scarabosio.